Ospreys director of coaching Scott Johnson is to leave the region at the end of the season and join the Scotland coaching set-up.
The 49-year-old Australian will become senior assistant coach to Scotland head coach Andy Robinson and will be in position for the June tour to Australia, Fiji and Samoa, the SRU has announced.
Johnson, a former Wales skills coach, Australia assistant coach and head coach of USA, was appointed as Ospreys director of coaching in April 2009.
He informed the board of his intention to move on ahead of Boxing Day's derby match against the Scarlets.
"Having initially committed myself to the Ospreys for three years, I've taken the time to consider my future plans and have advised the directors that while I intend to see out the remainder of my contract I will not be looking to renew at the end of the season," said Johnson.
"When I joined the region I had a remit to help set-up systems that would enable us to bring through home grown talent, coaches and players, a challenge I've thoroughly enjoyed getting my teeth into over the last three years.
"I feel that now is the right time for me to move on to other challenges but I remain 100 per cent committed to the Ospreys between now and the end of the season."
Johnson took charge of the Ospreys at the start of the 2009-10 season and the side reached the last eight of the Heineken Cup and won the first ever Magners League Grand Final.
Hugely missed
Ospreys Chief Operations Officer Andrew Hore said: "In 2008 we started a process which would see the Ospreys become a world leading player and coach development organisation with Scott joining us a year later as a part of this, to assist in producing such a development programme. Although this is now the end of the second phase, this process continues.
"As a region we have enjoyed success during his time with us but his biggest influence has been off the field, where he has played a major part in putting in place these new player and coach development structures while also having to deal with reducing the wage bill for the long-term good of the business. His work will help to ensure the long-term sustainability of the Ospreys."
"He'll be sadly missed by everybody at the Ospreys when he moves on, but by informing us of his intentions at this early stage, it allows us to use this time to consider our next course of action and the options available to us as we plan for the future."